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Ian Tizard

Distinguished Professor

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Phone: (979) 845-4276

Mailstop: 4467

Department: VTPB

Photo of Tizard, Ian


    Peru: The college has two formal agreements (Convenios) with the main Peruvian Veterinary Schools. Heredia Cayetana and San Marcos. Both commit us to collaborate in the area of avian research and conservation. There are multiple collaborative projects underway in Tambopata, in investigating the wild bird trade and in studying the epidemiology of selected avian diseases.

    Argentina: The Schubot Center is supporting a project to determine causes of nestling mortality in the Patagonian subspecies of the peregrine falcon.

    Canada: The Schubot Center is collaborating with scientists at the University of Guelph on the detection of avian bornavirus in arctic waterfowl.


  • Ph.D. Immunology, University of Cambridge, England 1969
  • B.Sc. Pathology, University of Edinburgh, Scotland, UK 1966
  • B.V.M.& S. Veterinary Medicine, University of Edinburgh, Scotland, UK 1965


  • ACVM (Honorary)
  • DSc (Honorary). University of Guelph

Scholarly Interests

Immunology with an emphasis on innovative vaccine technology and on the immunology of domestic mammals.Avian diseases with an emphasis on diseases of psittacinesPaleovirology,Genomics with an emphasis on whole avian genomes.The intestinal microbiome and its role in immunity.


Activation of a mouse macrophage cell line by acemannan: the major carbohydrate fraction from Aloe vera gel.
Zhang L, Tizard IR
Immunopharmacology. 35(2):119-28.
Acemannan, a beta-(1,4)-acetylated mannan, induces nitric oxide production in macrophage cell line RAW 264.7.
Ramamoorthy L, Kemp MC, Tizard IR
Mol Pharmacol. 50(4):878-84.
Antigen dependent adjuvant activity of a polydispersed beta-(1,4)-linked acetylated mannan (acemannan).
Chinnah AD, Baig MA, Tizard IR, Kemp MC
Vaccine. 10(8):551-7.
Aroclor 1254 as a 2,3,7,8-tetrachlorodibenzo-p-dioxin antagonist: effects on enzyme induction and immunotoxicity.
Bannister R, Davis D, Zacharewski T, Tizard I, Safe S
Toxicology. 46(1):29-42.
Development of a contagious ecthyma vaccine for goats.
Musser JM, Taylor CA, Guo J, Tizard IR, Walker JW
Am J Vet Res. 69(10):1366-70.
Positive reinforcement training facilitates the voluntary participation of laboratory macaws with veterinary procedures.
Daugette KF, Hoppes S, Tizard I, Brightsmith D
J Avian Med Surg. 26(4):248-54.
Proventricular dilatation disease in cockatiels (Nymphicus hollandicus) after infection with a genotype 2 avian bornavirus.
Mirhosseini N, Gray PL, Hoppes S, Tizard I, Shivaprasad HL, Payne S
J Avian Med Surg. 25(3):199-204.
Detection and characterization of a distinct bornavirus lineage from healthy Canada geese (Branta canadensis).
Payne S, Covaleda L, Jianhua G, Swafford S, Baroch J, Ferro PJ, Lupiani B, Heatley J, Tizard I
J Virol. 85(22):12053-6.
Comparison of sampling methods, culture, acid-fast stain, and polymerase chain reaction assay for the diagnosis of mycobacteriosis in ring-neck doves (Streptopelia risoria).
Saggese MD, Tizard I, Phalen DN
J Avian Med Surg. 24(4):263-71.
Molecular characterization of the cloacal microbiota of wild and captive parrots.
Xenoulis PG, Gray PL, Brightsmith D, Palculict B, Hoppes S, Steiner JM, Tizard I, Suchodolski JS
Vet Microbiol. 146(3-4):320-5.
Sickness behavior, its mechanisms and significance.
Tizard I
Anim Health Res Rev. 9(1):87-99.
The protective properties of milk and colostrum in non-human species.
Tizard I
Adv Nutr Res. 10:139-66.
Birds and bornaviruses.
Payne SL, Delnatte P, Guo J, Heatley JJ, Tizard I, Smith DA
Anim Health Res Rev. 13(2):145-56.
Grease, anthraxgate, and kennel cough: a revisionist history of early veterinary vaccines.
Tizard I
Adv Vet Med. 41:7-24.
Use of serologic testing to assess immune status of companion animals.
Tizard I, Ni Y
J Am Vet Med Assoc. 213(1):54-60.
Induction of apoptosis in a macrophage cell line RAW 264.7 by acemannan, a beta-(1,4)-acetylated mannan.
Ramamoorthy L, Tizard IR
Mol Pharmacol. 53(3):415-21.
Histopathology and the detection of avian bornavirus in the nervous system of birds diagnosed with proventricular dilatation disease.
Ouyang N, Storts R, Tian Y, Wigle W, Villanueva I, Mirhosseini N, Payne S, Gray P, Tizard I
Avian Pathol. 38(5):393-401.
Humoral response to Mycobacterium avium subsp. avium in naturally infected ring-neck doves (Streptopelia risoria).
Gray PL, Saggese MD, Phalen DN, Tizard I
Vet Immunol Immunopathol. 125(3-4):216-24.
Mycobacteriosis in naturally infected ring-neck doves (Streptopelia risoria): investigation of the association between feather colour and susceptibility to infection, disease and lesions type.
Saggese MD, Tizard I, Phalen DN
Avian Pathol. 37(4):443-50.
In situ detection and quantification of bursa of fabricius cellular proliferation or apoptosis in normal or steroid-treated neonatal chicks.
Higgins SE, Berghman LR, Moore RW, Caldwell DJ, Caldwell DY, Tizard I, Hargis BM
Poult Sci. 81(8):1136-41.
The biological activities of mannans and related complex carbohydrates.
Tizard IR, Carpenter RH, McAnalley BH, Kemp MC
Mol Biother. 1(6):290-6.
Evaluation of a killed, whole virion feline leukemia virus vaccine.
Tizard I, Bass EP
J Am Vet Med Assoc. 199(10):1410-3.
Characterization of a new genotype of avian bornavirus from wild ducks.
Guo J, Shivaprasad HL, Rech RR, Heatley JJ, Tizard I, Payne S
Virol J. 11(1):197.
Avian bornaviruses: diagnosis, isolation, and genotyping.
Guo J, Payne S, Zhang S, Turner D, Tizard I, Suchodolski P
Curr Protoc Microbiol. 34:15I.1.1-15I.1.33.
A draft de novo genome assembly for the northern bobwhite (Colinus virginianus) reveals evidence for a rapid decline in effective population size beginning in the Late Pleistocene.
Halley YA, Dowd SE, Decker JE, Seabury PM, Bhattarai E, Johnson CD, Rollins D, Tizard IR, Brightsmith DJ, Peterson MJ, Taylor JF, Seabury CM
PLoS One. 9(3):e90240.
Complete Genome Sequence of an Avian Bornavirus Isolated from a Healthy Canadian Goose (Branta canadensis).
Guo J, Baroch J, Randall A, Tizard I
Genome Announc. 1(5).
A multi-platform draft de novo genome assembly and comparative analysis for the Scarlet Macaw (Ara macao).
Seabury CM, Dowd SE, Seabury PM, Raudsepp T, Brightsmith DJ, Liboriussen P, Halley Y, Fisher CA, Owens E, Viswanathan G, Tizard IR
PLoS One. 8(5):e62415.
Ancient T-independence of mucosal IgX/A: gut microbiota unaffected by larval thymectomy in Xenopus laevis.
Mashoof S, Goodroe A, Du CC, Eubanks JO, Jacobs N, Steiner JM, Tizard I, Suchodolski JS, Criscitiello MF
Mucosal Immunol. 6(2):358-68.
Bioencapsulation of the hepatitis B surface antigen and its use as an effective oral immunogen.
Hayden CA, Streatfield SJ, Lamphear BJ, Fake GM, Keener TK, Walker JH, Clements JD, Turner DD, Tizard IR, Howard JA
Vaccine. 30(19):2937-42.
Intranasal delivery of Norwalk virus-like particles formulated in an in situ gelling, dry powder vaccine.
Velasquez LS, Shira S, Berta AN, Kilbourne J, Medi BM, Tizard I, Ni Y, Arntzen CJ, Herbst-Kralovetz MM
Vaccine. 29(32):5221-31.
The diagnosis of proventricular dilatation disease: use of a Western blot assay to detect antibodies against avian Borna virus.
Villanueva I, Gray P, Mirhosseini N, Payne S, Hoppes S, Honkavuori KS, Briese T, Turner D, Tizard I
Vet Microbiol. 143(2-4):196-201.
Stabilization of growth factors relevant to wound healing by a plant cell wall biomaterial.
Ni Y, Turner D, Yates K, Tizard I
Planta Med. 73(12):1260-6.
Supercritical fluid extraction provides an enhancement to the immune response for orally-delivered hepatitis B surface antigen.
Hayden CA, Smith EM, Turner DD, Keener TK, Wong JC, Walker JH, Tizard IR, Jimenez-Flores R, Howard JA
Vaccine. 32(11):1240-6.
Isolation and characterization of structural components of Aloe vera L. leaf pulp.
Ni Y, Turner D, Yates KM, Tizard I
Int Immunopharmacol. 4(14):1745-55.
Specificity and prevalence of natural bovine antimannan antibodies.
Srinivasan A, Ni Y, Tizard I
Clin Diagn Lab Immunol. 6(6):946-52.
Use of immunomodulators as an aid to clinical management of feline leukemia virus-infected cats.
Tizard I
J Am Vet Med Assoc. 199(10):1482-5.
Risks associated with use of live vaccines.
Tizard I
J Am Vet Med Assoc. 196(11):1851-8.
Avian bornavirus and proventricular dilatation disease: diagnostics, pathology, prevalence, and control.
Hoppes SM, Tizard I, Shivaprasad HL
Vet Clin North Am Exot Anim Pract. 16(2):339-55.

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